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  <url>
    <loc>https://www.santarosaprimates.org/home</loc>
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    <lastmod>2023-11-07</lastmod>
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      <image:title>Santa Rosa Primate Project</image:title>
    </image:image>
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      <image:title>Santa Rosa Primate Project</image:title>
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      <image:title>Santa Rosa Primate Project</image:title>
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  <url>
    <loc>https://www.santarosaprimates.org/santarosaprimateproject/about</loc>
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    <priority>0.75</priority>
    <lastmod>2023-10-05</lastmod>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/9efaf9aa-455a-48a6-88e6-8d66a076a2f7/map+full+size.jpeg</image:loc>
      <image:title>About - A Biodiversity Hotspot</image:title>
      <image:caption>We work in a place of incredible biodiversity in the Área de Conservación Guanacaste, Costa Rica (ACG). The ACG has a fascinating history and stands as an incredible conservation success story. The ACG harbors an outstanding 2.6% of terrestrial global biodiversity and stands as the largest tract of critically endangered tropical dry forest in the world–connecting coastal forests to dry forests to rainforests and cloud forests. Our project is based in the Santa Rosa Sector of the ACG. This sector offers a unique setting for study due to its status as a regenerating, secondary growth, tropical dry forest. This has provided us a rare opportunity to assess effects of forest protection and regeneration on resident primate species.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/e38dac57-04d1-4a21-b3f8-d1dfe436f81d/69719938-710b-4947-904e-e5ff95eb564d.JPG</image:loc>
      <image:title>About</image:title>
      <image:caption>Young capuchins playing. Photo by Nick Chapoy.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/3198da63-08c6-440b-b6f5-cc78e4258f62/howler.jpeg</image:loc>
      <image:title>About</image:title>
      <image:caption>Howler money (Allouatta palliata). Photo by Nick Chapoy.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/e42bc132-4754-4f55-8840-f0c027f56471/trees+small.jpeg</image:loc>
      <image:title>About</image:title>
      <image:caption>Santa Rosa secondary growth forest in the wet season. Photo by Nelle Kulick.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1663774176217-18S85YYV1MEQ4STP2SQA/Campos-2006-06-26_06-.jpg</image:loc>
      <image:title>About</image:title>
      <image:caption>View from the historical monument at the Casona overlooking Santa Rosa and other sectors of the ACG. Photo by Fernando Campos.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1663774581436-5YC55E1XXLGPMNFSOXNN/Campos-2006-05-15_16-.jpg</image:loc>
      <image:title>About</image:title>
      <image:caption>Santa Rosa during the wet/dry season transition. Photo by Fernando Campos.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1663776250824-SLIHY0QX1GJY6M5TE4KD/Campos-20100306-IMG_5376-.jpg</image:loc>
      <image:title>About</image:title>
      <image:caption>Firebreak and patch of early successional forest in Santa Rosa in the dry season. Photo by Fernando Campos.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/c9144bfe-3286-4742-98b1-09b5a50c2b91/marcos+spider.jpeg</image:loc>
      <image:title>About</image:title>
      <image:caption>Spider monkey mother and infant. Photo by Marcos Alvarez Garita.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1663776130019-IMR7DF7GG1F7LAJCJV13/Campos-20100216-IMG_5135-.jpg</image:loc>
      <image:title>About</image:title>
      <image:caption>Santa Rosa during the wet/dry season transition. Photo by Fernando Campos.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/fc72260a-e8ac-41bc-9a3e-5542e6ed86c0/NICK+SPIDER.jpeg</image:loc>
      <image:title>About</image:title>
      <image:caption>Spider Monkey (Ateles geoffroyi). Photo by Nick Chapoy.</image:caption>
    </image:image>
  </url>
  <url>
    <loc>https://www.santarosaprimates.org/contact</loc>
    <changefreq>daily</changefreq>
    <priority>0.75</priority>
    <lastmod>2022-09-22</lastmod>
  </url>
  <url>
    <loc>https://www.santarosaprimates.org/our-team</loc>
    <changefreq>daily</changefreq>
    <priority>0.75</priority>
    <lastmod>2025-06-20</lastmod>
    <image:image>
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      <image:title>Our Team</image:title>
    </image:image>
    <image:image>
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      <image:title>Our Team</image:title>
    </image:image>
    <image:image>
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      <image:title>Our Team</image:title>
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    <image:image>
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      <image:title>Our Team</image:title>
    </image:image>
    <image:image>
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      <image:title>Our Team</image:title>
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      <image:title>Our Team</image:title>
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    <image:image>
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      <image:title>Our Team</image:title>
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      <image:title>Our Team</image:title>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/f5a7e000-e708-4ff6-9e03-3992df07bf36/shasta+team.jpeg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Team Microbiome, 2022. Left to right: Megan Mah, Alice Poirer, Shasta Webb, Michelle Reed</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656920997677-UX51YHIIM573LT581XM0/su%2Band%2Bamanda.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Suheidy Romero Morales (left) and Dr. Amanda Melin (right) logging samples</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1665504941143-HCRHXAJGOL7W42SS2M0S/296591113_785312269196840_3123765420352275926_n.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Julia Casorso videotaping foraging behavior in the field</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921042815-3XPW3Y0BK4AEG75OBBG9/allegra%2Bmegan%2Bx.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Allegra DePasquale (front left), Megan Mah (back left) and Cynthia (right) collecting plant samples to collect data on fruit and pith traits</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921174321-5OEWKUFBTHWXEYTDNEFV/02cee316-dded-4437-aef4-6cd28bdf8cfc.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Saúl Cheves (left) and Catalina Chaves-Cordero (right) processing samples for DNA sequencing</image:caption>
    </image:image>
    <image:image>
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      <image:title>Our Team</image:title>
      <image:caption>2019 Field Team participating in an event with ¡Quiero Dejar Una Huella Verde!</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921017006-JWN3074C18LW1XB9LH8X/c1c03046-94a4-41c8-afc6-71a47685ad49%2B2.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Nelle Kulick (left) and Dr. Katharine Jack (right) collecting urine samples for chemical analysis</image:caption>
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    <image:image>
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      <image:title>Our Team</image:title>
      <image:caption>Alice Poirer's first urine sample collection on Odor Team. Go Alice!</image:caption>
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    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921133493-VY450JV6FPE9HYM3HMAS/IMG_2141.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Team excited for our 2022 park-wide population census of capuchin and howler monkeys</image:caption>
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    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656920979180-TUXGE3NOJQQECFGTIV4W/team.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Odor Team ready for the day! From left to right, front to back: Marcos Alvarez Garita, Alice Poirer, Suheidy Romero Morales, Giulia Severino, Jonathan Wilkins, Yarenis Abella</image:caption>
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    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921062845-V1NS8YT0W0VVSAN7UBQU/saul%2Bronald%2Bsu.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>All smiles after a long day in the forest! Left to right: Saúl, Ronald, and Suheidy</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921314477-Y658HUG4D5CNNBCRT0JZ/8A967879-56D9-4121-AF69-D7989D69DB0B.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Margaret Buehler running ELISAs in the field laboratory at the Capuchinos de Taboga Project. Thank you for having us!</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1657073748169-0FF0N2DWZFPGBWBMDAV3/su%2Bteaching%2Bdani%2B5.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Suheidy teaching urine sample collection methods in the field</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921085624-F4JSELD2DNOW6QHKAZUO/IMG_4910.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Marcos Alvarez Garita giving a presentation on our work to a class of local university students</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1665503834752-OBV8UF8M4YWYJ8XUBCB9/309745444_779470809983362_6012776257435348735_n.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Julia Casorso using the mechanical tester to measure fruit traits</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921196237-O4YYI4JAC5WBS1X77237/a6145419-065d-41a4-a10d-f7e98cd18db2.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Nick Chapoy with his first collected urine sample on Odor Team. Way to go, Nick!</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1665506870762-Q2VGEBQKN3X79O7SIDY5/nelle%252Bcat%252Bgabe.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Team Chemical Communication, 2021. Left to right: Catalina Chaves-Cordero, Nelle Kulick, Gabe Benson.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921334401-CHJKT73NSM0QEL0V04YX/3539bb60-261c-41c8-a785-40ace68d5a6d.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Yarenis Abella updating sample logs after performing fecal hormone extractions in the lab</image:caption>
    </image:image>
    <image:image>
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      <image:title>Our Team</image:title>
      <image:caption>Suheidy, Nick, and Wil calibrating the laser box we use to non-invasively measure body parts of alpha and subordinate male capuchins from photographs</image:caption>
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    <image:image>
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      <image:title>Our Team</image:title>
      <image:caption>Team Foraging and Color Vision, 2019. Left to right: JC, Allegra, Megan, and Cynthia</image:caption>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921220002-84K0GSTI5UMUIXEP7HF9/IMG_4905.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Dani Salazar collecting a urine sample in the field. Awesome work, Dani!</image:caption>
    </image:image>
    <image:image>
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      <image:title>Our Team</image:title>
      <image:caption>The team was all smiles as Marcos collected the first urine sample of the season!</image:caption>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/a7189d4d-9e63-4c1a-99f3-e8d5c2bc34fd/great+bino+pic.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Team practicing monkey identifications using natural markings (e.g., differences in fur and skin colorations, scars, broken digits)</image:caption>
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      <image:title>Our Team</image:title>
      <image:caption>Odor Team 2022. Left to right: Nick Chapoy, Nelle Kulick, Marcos Alvarez Garita, Dr. Katharine Jack, Giulia Severino, Suheidy Romero Morales</image:caption>
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      <image:title>Our Team</image:title>
      <image:caption>Saúl taking a break while searching for monkeys, Census 2022</image:caption>
    </image:image>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1665503787915-QCG9HAXWDJ0JBRENNSLI/308711510_804454160699572_5421540503611544244_n.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Julia Casorso using photogrammetry methods to collect data on foraging behavior</image:caption>
    </image:image>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1656921366205-SSEN4LLSBAB4J9AZ9SOI/488572ac-a7f0-4556-984a-5c6267de7de0.jpg</image:loc>
      <image:title>Our Team</image:title>
      <image:caption>Dr. Katharine Jack teaching urine sample collection methods in the field</image:caption>
    </image:image>
  </url>
  <url>
    <loc>https://www.santarosaprimates.org/research</loc>
    <changefreq>daily</changefreq>
    <priority>0.75</priority>
    <lastmod>2023-08-06</lastmod>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/3581b5bf-4db4-4bf4-a9c6-79bdd6c1337a/Legolas-scaled.jpg</image:loc>
      <image:title>Research - Make it stand out</image:title>
      <image:caption>At almost 30, Legolas was one of the oldest individuals in our study population</image:caption>
    </image:image>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/2c53d335-fd0e-45a3-9b44-85e3b495dcdb/2593fbe5-5bea-4c67-9c00-95f060035bf5.JPG</image:loc>
      <image:title>Research</image:title>
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    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/197f24e5-b53a-46cd-a7a0-0598d0751f1d/image-asset+%281%29.jpeg</image:loc>
      <image:title>Research - Sensory and Foraging Ecology</image:title>
      <image:caption>Principal Investigator: Dr. Amanda Melin Through their senses, animals interface with the environment to find food, select mates, and avoid predators. The interrelationships among diet, activity pattern and the senses are central to hypotheses of primate origins and adaptive radiation and shape the ecological niches of extant species. Through the integration of observational fieldwork, genetics and genomics, and computer models of animal vision, we seek to understand the adaptive complexes of primates, including humans, using an innovative and multidisciplinary approach. Our research centers around: 1) intra- and interspecific color vision variation in primates and other mammals with functional and phylogenetic relevance to primate origins and adaptive radiation (treeshrews, fruit bats, opossums); 2) variation within and between species in diet and use of sensory behaviors (e.g. touch, sniff, lick) while foraging, with the goal of linking behavioral variation to sensory genotypes; and 3) the visual, olfactory, gustatory and mechanical cues of food quality, with the goal of identifying important cues that shape the sensory systems of consumers.</image:caption>
    </image:image>
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      <image:title>Research - Make it stand out</image:title>
      <image:caption>Whatever it is, the way you tell your story online can make all the difference.</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/4166d282-8b84-4a00-8ded-25127e006970/eating+insects.jpg</image:loc>
      <image:title>Research - Plasticity and Adaptation in the Gut Microbiome</image:title>
      <image:caption>Principal Investigator: Dr. Amanda Melin Recent research on the human microbiome has demonstrated a strong effect of diet, environment and health on the gut microbiota, raising the possibility that this relationship extends to other primates. However, little is known about how changing environments, accompanied by pronounced dietary shifts, affects the microbiome and shapes digestive adaptations. Ongoing advances in massive parallel sequencing are continually increasing our ability to ask refined, detailed question of wild populations. We are conducting a fecal metagenomic study spanning the seasonal transitions in Santa Rosa, Costa Rica to identify the taxonomic composition and functional genomes of the resident intestinal microbiota of capuchin monkeys and reveal how shifts in climate and diet impact these symbiotic organisms. We are additionally assembling a genone reference for white-faced capuchin monkeys that will assist in this project and be broadly useful in comparative primate genomics. Together, this information will reveal how omnivorous primates manage the varied digestive challenges of their eclectic diet, and adapt to the profound seasonal shifts in food availability.</image:caption>
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      <image:title>Research - Making an Alpha Male: the Behavior, Endocrinology, and Genetics of Dominance in Male White-faced Capuchins</image:title>
      <image:caption>Principal Investigator: Dr. Katharine Jack We are examining behavioral, endocrinological, and genetic correlates (including kinship, reproductive success, and the major histocompatibility complex [MHC]) of male life history and reproductive strategies. The main goal of this longitudinal project is to understand how hormonal and social factors influence the emergence, from puberty into adulthood, of dominance status in male white-faced capuchins (e.g. Jack et al., 2014). Identifying traits and processes influencing individual variability in reproductive success is fundamental to understanding evolutionary processes. In our capuchins, like many other socially living mammals including humans, there is a strong association between dominance status and reproductive success, however, we still know very little about how and why some males attain alpha status while others do not. These data, along with our ongoing studies of the role of the major histocompatibility in male dispersal and mate choice decisions (funded by the LA Board of Regents, with Jessica Lynch Alfaro), will provide us with a holistic picture for examining male dominance and reproductive biology.</image:caption>
    </image:image>
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      <image:caption>Whatever it is, the way you tell your story online can make all the difference.</image:caption>
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    <lastmod>2022-06-21</lastmod>
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    <loc>https://www.santarosaprimates.org/conservation-education-and-outreach</loc>
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    <lastmod>2022-09-22</lastmod>
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      <image:title>Conservation, Education, and Outreach</image:title>
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      <image:title>Conservation, Education, and Outreach</image:title>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/5ec321c2af33de48734cc929/1607694644871-IC85FNH781UNZSZEGHDR/Aro+Ha_0428.jpg</image:loc>
      <image:title>Conservation, Education, and Outreach</image:title>
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      <image:title>Conservation, Education, and Outreach</image:title>
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    <loc>https://www.santarosaprimates.org/local-training</loc>
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    <lastmod>2022-09-21</lastmod>
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      <image:caption>Performing hormone extractions</image:caption>
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      <image:title>Local Training</image:title>
      <image:caption>Collecting biological samples in the field</image:caption>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1657080825093-1JIZJOXE5ENNFMN9G7X6/8f510cb3-1a9b-4988-b8d6-dbe8e73e77a3.jpg</image:loc>
      <image:title>Local Training</image:title>
      <image:caption>Working in the lab</image:caption>
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    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1657080937338-FMB6RFGL2R9UOMCJXGRF/su%2Band%2Bjulia%2B2.jpg</image:loc>
      <image:title>Local Training</image:title>
      <image:caption>Collecting behavioral data</image:caption>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1657080953409-8O7K3SS48AEC4DW81YDX/3539bb60-261c-41c8-a785-40ace68d5a6d.jpg</image:loc>
      <image:title>Local Training</image:title>
      <image:caption>Logging biological samples</image:caption>
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      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1665504625328-M0WRRSATWREH2KHCDCWP/309593489_1099011307422125_6603082798457499508_n.jpg</image:loc>
      <image:title>Local Training</image:title>
      <image:caption>Collecting data on foraging behavior using photogrammetry</image:caption>
    </image:image>
    <image:image>
      <image:loc>https://images.squarespace-cdn.com/content/v1/62966ff0001e3542e76acdb9/1665504439524-RAQOTJG44324VBWWMDE5/309747673_5458139460929602_6071508237329239028_n.jpg</image:loc>
      <image:title>Local Training</image:title>
      <image:caption>Collecting data on fruit traits using a mechanical tester</image:caption>
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    <loc>https://www.santarosaprimates.org/publications</loc>
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    <lastmod>2024-10-14</lastmod>
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      <image:caption>N. Chapoy</image:caption>
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      <image:caption>F. Campos</image:caption>
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      <image:caption>N. Kulick</image:caption>
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      <image:caption>A. Melin</image:caption>
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      <image:caption>N. Kulick</image:caption>
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      <image:caption>N. Chapoy</image:caption>
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      <image:caption>N. Chapoy</image:caption>
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      <image:title>Photo Gallery</image:title>
      <image:caption>N. Kulick</image:caption>
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      <image:caption>F. Campos</image:caption>
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      <image:caption>N. Chapoy</image:caption>
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      <image:caption>N. Kulick</image:caption>
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      <image:caption>A . Melin</image:caption>
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      <image:caption>C. Martinez</image:caption>
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      <image:caption>C. Martinez</image:caption>
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      <image:caption>F. Campos</image:caption>
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      <image:caption>N. Kulick</image:caption>
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      <image:caption>F. Campos</image:caption>
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      <image:caption>N. Chapoy</image:caption>
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